Originally posted by yunger
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On my first suggestion I mentioned the cap, but as Enzo pointed out I hadn't seen C6 on the upper left corner. So that stage is supposedly decoupled, assuming those caps are good(like I said(half jokingly) and JMFahey too, I'd add another resistor there anyway just before the first plate resistor). That 12ax7 will not suffer at all from some more added resistance.
The 12at7 is in phase with your first gain stage when you have 2 gain stages(full circuit), so when you jumped over the second gain stage you set the preamp and power amp to be out of phase. If that 12at7 is modulating the power supply, that'd be a plausible explanation for your motorboating. Again, two small tubes should not be able to mess with your power supply. I made a tiny 12ax7 + 12au7 amp with just 20uF reservoir and 10uF for second filter....no oscillations of any kind....so like others said, there is enough capacitance in there to keep it stable.
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