I think shorting it out makes the reverb tank output transducer almost short that V1B grid to ground.
...any cap differences can lead to phase cancellations..
Don't know why they choose this driver circuit. But the 2 triodes are actually wired in parallel for AC signals. Maybe the reverb transformer was not suitable for primary DC current (no air-gap) and/ or connecting the plates would have caused instabilities.
Bypassing R42 will increase reverb return level.
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